本文主要研究内容
作者王茁（2019）在《玫瑰黄腹三节叶蜂幼虫侧单眼形态和超微结构》一文中研究指出：侧单眼stemmata是全变态类幼虫特有的视觉器官,一般是由角膜晶体、视网膜细胞、色素细胞、角膜生成细胞以及晶体等结构组成。然而,全变态类不同类群幼虫的侧单眼,无论是数目还是内部结构方面,都存在着较大的差异,甚至同一种幼虫的不同侧单眼在结构方面都存在较大差异。鳞翅目Lepidoptera、鞘翅目Coleoptera、广翅目Megaloptera等幼虫侧单眼已进行了较多的研究,但仅广腰亚目Symphyta幼虫存在1对侧单眼的膜翅目Hymenoptera中研究较少,目前仅报道了筒腹叶蜂科Pergidae和叶蜂科Tenthredinidae幼虫侧单眼的结构,而三节叶蜂科Argidae幼虫侧单眼的结构迄今未见报道。本文利用光学显微镜、扫描电子显微镜和透射电子显微镜,研究了玫瑰黄腹三节叶蜂Arge pagana(Panzer,1798)(膜翅目:三节叶蜂科)幼虫侧单眼的外部形态和内部结构,并依据侧单眼结构推测其可能的功能,讨论了全变态类幼虫侧单眼的起源和进化。玫瑰黄腹三节叶蜂幼虫侧单眼主要是由角膜晶体、视网膜细胞、色素细胞和角膜生成细胞等结构组成,无晶体。该角膜晶体透明且强烈双凸,表面具有许多浅浅的凹陷;角膜晶体外缘被一个黑色圆环包围,该圆环的后缘有缺刻。角膜生成细胞位于角膜晶体的下方,其远端部分包含了大量囊泡并且形成一个相对电子透明的区域,其近端部分延伸至基膜附近。视网膜细胞位于透明区域的下方。该侧单眼包含了大约1000个视网膜细胞,其中相邻的8个或9个视网膜细胞聚成一簇,一共形成大约100个视杆。由8个视网膜细胞形成的视杆和由9个视网膜细胞形成的视杆位于2个不同区域,区域I和区域II。区域I位于侧单眼的中央占大多数面积,视杆由8个视网膜细胞形成。区域II位于侧单眼的后缘呈条带分布,视杆由9个视网膜细胞形成。在该侧单眼的最后缘,即区域II的外侧,分布着色素细胞束;这些色素细胞束从角膜生成细胞的远端部延伸至视网膜细胞的近端部。由于玫瑰黄腹三节叶蜂幼虫侧单眼的后缘存在少量由9个视网膜细胞形成的视杆,这些视杆的视小杆大致垂直排列,由此推测玫瑰黄腹三节叶蜂幼虫侧单眼可能对偏振光具有敏感性。玫瑰黄腹三节叶蜂幼虫侧单眼与筒腹叶蜂科幼虫侧单眼结构相似,都是由角膜晶体、角膜生成细胞和视网膜细胞组成,无晶体。其中,筒腹叶蜂科幼虫侧单眼被认为起源进化于多个小眼角膜的融合和晶体的退化,但其视杆仍由8个视网膜细胞形成保持原始状态。本研究发现,玫瑰黄腹三节叶蜂幼虫侧单眼内的视杆由8或9个视网膜细胞形成;由8个视网膜细胞形成的视杆位于中央,占大多数,由9个视网膜细胞形成的视杆位于后缘,占少数,因此推测玫瑰黄腹三节叶蜂幼虫侧单眼可能进化起源于两种类型的小眼融合。
Abstract
ce chan yan stemmatashi quan bian tai lei you chong te you de shi jiao qi guan ,yi ban shi you jiao mo jing ti 、shi wang mo xi bao 、se su xi bao 、jiao mo sheng cheng xi bao yi ji jing ti deng jie gou zu cheng 。ran er ,quan bian tai lei bu tong lei qun you chong de ce chan yan ,mo lun shi shu mu hai shi nei bu jie gou fang mian ,dou cun zai zhao jiao da de cha yi ,shen zhi tong yi chong you chong de bu tong ce chan yan zai jie gou fang mian dou cun zai jiao da cha yi 。lin chi mu Lepidoptera、qiao chi mu Coleoptera、an chi mu Megalopteradeng you chong ce chan yan yi jin hang le jiao duo de yan jiu ,dan jin an yao ya mu Symphytayou chong cun zai 1dui ce chan yan de mo chi mu Hymenopterazhong yan jiu jiao shao ,mu qian jin bao dao le tong fu xie feng ke Pergidaehe xie feng ke Tenthredinidaeyou chong ce chan yan de jie gou ,er san jie xie feng ke Argidaeyou chong ce chan yan de jie gou qi jin wei jian bao dao 。ben wen li yong guang xue xian wei jing 、sao miao dian zi xian wei jing he tou she dian zi xian wei jing ,yan jiu le mei gui huang fu san jie xie feng Arge pagana(Panzer,1798)(mo chi mu :san jie xie feng ke )you chong ce chan yan de wai bu xing tai he nei bu jie gou ,bing yi ju ce chan yan jie gou tui ce ji ke neng de gong neng ,tao lun le quan bian tai lei you chong ce chan yan de qi yuan he jin hua 。mei gui huang fu san jie xie feng you chong ce chan yan zhu yao shi you jiao mo jing ti 、shi wang mo xi bao 、se su xi bao he jiao mo sheng cheng xi bao deng jie gou zu cheng ,mo jing ti 。gai jiao mo jing ti tou ming ju jiang lie shuang tu ,biao mian ju you hu duo jian jian de ao xian ;jiao mo jing ti wai yuan bei yi ge hei se yuan huan bao wei ,gai yuan huan de hou yuan you que ke 。jiao mo sheng cheng xi bao wei yu jiao mo jing ti de xia fang ,ji yuan duan bu fen bao han le da liang nang pao bing ju xing cheng yi ge xiang dui dian zi tou ming de ou yu ,ji jin duan bu fen yan shen zhi ji mo fu jin 。shi wang mo xi bao wei yu tou ming ou yu de xia fang 。gai ce chan yan bao han le da yao 1000ge shi wang mo xi bao ,ji zhong xiang lin de 8ge huo 9ge shi wang mo xi bao ju cheng yi cu ,yi gong xing cheng da yao 100ge shi gan 。you 8ge shi wang mo xi bao xing cheng de shi gan he you 9ge shi wang mo xi bao xing cheng de shi gan wei yu 2ge bu tong ou yu ,ou yu Ihe ou yu II。ou yu Iwei yu ce chan yan de zhong yang zhan da duo shu mian ji ,shi gan you 8ge shi wang mo xi bao xing cheng 。ou yu IIwei yu ce chan yan de hou yuan cheng tiao dai fen bu ,shi gan you 9ge shi wang mo xi bao xing cheng 。zai gai ce chan yan de zui hou yuan ,ji ou yu IIde wai ce ,fen bu zhao se su xi bao shu ;zhe xie se su xi bao shu cong jiao mo sheng cheng xi bao de yuan duan bu yan shen zhi shi wang mo xi bao de jin duan bu 。you yu mei gui huang fu san jie xie feng you chong ce chan yan de hou yuan cun zai shao liang you 9ge shi wang mo xi bao xing cheng de shi gan ,zhe xie shi gan de shi xiao gan da zhi chui zhi pai lie ,you ci tui ce mei gui huang fu san jie xie feng you chong ce chan yan ke neng dui pian zhen guang ju you min gan xing 。mei gui huang fu san jie xie feng you chong ce chan yan yu tong fu xie feng ke you chong ce chan yan jie gou xiang shi ,dou shi you jiao mo jing ti 、jiao mo sheng cheng xi bao he shi wang mo xi bao zu cheng ,mo jing ti 。ji zhong ,tong fu xie feng ke you chong ce chan yan bei ren wei qi yuan jin hua yu duo ge xiao yan jiao mo de rong ge he jing ti de tui hua ,dan ji shi gan reng you 8ge shi wang mo xi bao xing cheng bao chi yuan shi zhuang tai 。ben yan jiu fa xian ,mei gui huang fu san jie xie feng you chong ce chan yan nei de shi gan you 8huo 9ge shi wang mo xi bao xing cheng ;you 8ge shi wang mo xi bao xing cheng de shi gan wei yu zhong yang ,zhan da duo shu ,you 9ge shi wang mo xi bao xing cheng de shi gan wei yu hou yuan ,zhan shao shu ,yin ci tui ce mei gui huang fu san jie xie feng you chong ce chan yan ke neng jin hua qi yuan yu liang chong lei xing de xiao yan rong ge 。
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论文作者分别是来自西北农林科技大学的王茁，发表于刊物西北农林科技大学2019-07-11论文，是一篇关于全变态类论文,视觉器官论文,视杆论文,偏振光论文,进化论文,西北农林科技大学2019-07-11论文的文章。本文可供学术参考使用，各位学者可以免费参考阅读下载，文章观点不代表本站观点，资料来自西北农林科技大学2019-07-11论文网站，若本站收录的文献无意侵犯了您的著作版权，请联系我们删除。